First, that looks great and can we have a close up of the fenders?

I'd say level the saddle for one and just give it a shot. It may be standard to tilt it upward - but it's your bum & other parts that are on it, so do what feels good for you.

Handlebars that sweep further back towards you might help.

Something that sometimes helps me is putting a longer higher angle stem on bikes, so maybe try different stem angles and see if it sits you more upright.

You can always switch saddles on the bikes for a little bit and see if you like the geometry better with your usual saddle.