I adjusted my saddle from a slight upward tilt to netural even. I was incredibly amazed at the result. Such a minor adjustment made such a big difference. One thing I noticed was that the change also changed weight distribution. I found with the neutral setting that I was now putting more weight on my forearms, shoulders, wrists and hands on the handlebars. The seat was set to the correct height so I raised the handlebars in micro increments (like 1/8 and 1/4 of an inch) until I got it just perfect.

You are wise to go on a longer ride and continue to tweak until you get it goldielocks "just right".