nscrbug, have you tried running with the injinji toe socks? My feet look a lot like yours, though running has never been all that painful for me. My biggest issue was that the toenails on the curled-in toes would routinely cut holes in the toes beside them on long runs. Several pairs of bloody socks later, I bought a pair of the injinjis just to see if forcing them to separate would help. Definitely helped the nail cuts, but maybe it would help your pain by forcing those toes to spread a bit? Just an idea.